I don't think we know a whole lot yet, and there is certainly extremely little or no information from there on anything downstream of M269.Cinnioglu's 2004 paper, Excavating Y Chromosome Haplotype Strata in Anatolia
, remains the most comprehensive work on the subject, I believe.
In the last year or so word has come out of Russia from the university at Ufa in Bashkortostan of the work of Khusnutdinova and Yunusbaev and others among the Bashkirs that reveals that the Bashkirs of the Volga-Ural region have a high frequency of R-M269. Overall the average is 50%, and in someplaces it is as high as 82%.
But we don't know about anything downstream of M269.
A couple of Bashkirs are suppsoed to have submitted kits to FTDNA, but I haven't received any word of their results yet.